Hmmm... There was something dodgy going on. Scoping the input stage after half an hour's power up time revealed intermittent bursts of 50Hz hum, mixed with higher frequency spiky stuff appearing at the cathode of the left hand valve, with the same kind of dirty waveform bursts at the right hand valve, but at a lower amplitude.I'm going to have to investigate voltages and currents around the circuit, to be sure it is not something untoward going on.
Further physical investigation revealed a dodgy looking situation between one of the 4V heater legs and the cathode tag of the left valve, where there was a double heater connection to carry the 4V over to the right valve.
Also the left hand valve grid connection wire from the volume control was trapped under one of the legs of the heater wiring.
Solder sucking the heater and grid tags of the left hand valve socket and rerouting the offending grid wire, seems (touch wood) to have cured the noise. The amp has been idling with 41MTL mesh plates in situ for 45 minutes with no further bursts of dirty noise from the speakers.
It is probably now possible to say that there is nothing wrong with either of the 41MTLs and by inference probably nothing amiss with the MH4s either.
The fault looks like it was either an intermittent cathode to heater or grid to heater short at the socket.
"Job's a guddun", as they say.